Regulatory sites for splicing in human basal ganglia are enriched for disease-relevant information


Por: Guelfi S., D’Sa K., Botía J.A., Vandrovcova J., Reynolds R.H., Zhang D., Trabzuni D., Collado-Torres L., Thomason A., Quijada Leyton P., Gagliano Taliun S.A., Nalls M.A., Noyce A.J., Nicolas A., Cookson M.R., Bandres-Ciga S., Gibbs J.R., Hernandez D.G., Singleton A.B., Reed X., Leonard H., Blauwendraat C., Faghri F., Bras J., Guerreiro R., Tucci A., Kia D.A., Houlden H., Plun-Favreau H., Mok K.Y., Wood N.W., Lovering R., R’Bibo L., Rizig M., Chelban V., Tan M., Morris H.R., Middlehurst B., Quinn J., Billingsley K., Holmans P., Kinghorn K.J., Lewis P., Escott-Price V., Williams N., Foltynie T., Brice A., Danjou F., Lesage S., Corvol J.-C., Martinez M., Giri A., Schulte C., Brockmann K., Simón-Sánchez J., Heutink P., Gasser T., Rizzu P., Sharma M., Shulman J.M., Robak L., Lubbe S., Mencacci N.E., Finkbeiner S., Lungu C., Scholz S.W., Gan-Or Z., Rouleau G.A., Krohan L., van Hilten J.J., Marinus J., Adarmes-Gómez A.D., Bernal-Bernal I., Bonilla-Toribio M., Buiza-Rueda D., Carrillo F., Carrión-Claro M., Mir P., Gómez-Garre P., Jesús S., Labrador-Espinosa M.A., Macias D., Vargas-González L., Méndez-del-Barrio C., Periñán-Tocino T., Tejera-Parrado C., Diez-Fairen M., Aguilar M., Alvarez I., Boungiorno M.T., Carcel M., Pastor P., Tartari J.P., Alvarez V., González M.M., Blazquez M., Garcia C., Suarez-Sanmartin E., Barrero F.J., Rezola E.M., Yarza J.A.B., Pagola A.G., Arregui A.L.M., Ruiz-Martínez J., Cerdan D., Duarte J., Clarimón J., Dols-Icardo O., Infante J., Marín J., Kulisevsky J., Pagonabarraga J., Gonzalez-Aramburu I., Rodriguez A.S., Sierra M., Duran R., Ruz C., Vives F., Escamilla-Sevilla F., Mínguez A., Cámara A., Compta Y., Ezquerra M., Marti M.J., Fernández M., Muñoz E., Fernández-Santiago R., Tolosa E., Valldeoriola F., García-Ruiz P., Heredia M.J.G., Errazquin F.P., Hoenicka J., Jimenez-Escrig A., Martínez-Castrillo J.C., Lopez-Sendon J.L., Torres I.M., Tabernero C., Vela L., Zimprich A., Pihlstrom L., Koks S., Taba P., Majamaa K., Siitonen A., Okubadejo N.U., Ojo O.O., Forabosco P., Walker R., Small K.S., Smith C., Ramasamy A., Hardy J., Weale M.E., Ryten M.

Publicada: 1 ene 2020
Resumen:
Genome-wide association studies have generated an increasing number of common genetic variants associated with neurological and psychiatric disease risk. An improved understanding of the genetic control of gene expression in human brain is vital considering this is the likely modus operandum for many causal variants. However, human brain sampling complexities limit the explanatory power of brain-related expression quantitative trait loci (eQTL) and allele-specific expression (ASE) signals. We address this, using paired genomic and transcriptomic data from putamen and substantia nigra from 117 human brains, interrogating regulation at different RNA processing stages and uncovering novel transcripts. We identify disease-relevant regulatory loci, find that splicing eQTLs are enriched for regulatory information of neuron-specific genes, that ASEs provide cell-specific regulatory information with evidence for cellular specificity, and that incomplete annotation of the brain transcriptome limits interpretation of risk loci for neuropsychiatric disease. This resource of regulatory data is accessible through our web server, http://braineacv2.inf.um.es/. © 2020, The Author(s).

Filiaciones:
Guelfi S.:
 Reta Lila Weston Research Laboratories, Department of Molecular Neuroscience, University College London (UCL) Institute of Neurology, London, United Kingdom

D’Sa K.:
 Reta Lila Weston Research Laboratories, Department of Molecular Neuroscience, University College London (UCL) Institute of Neurology, London, United Kingdom

 Department of Medical & Molecular Genetics, School of Medical Sciences, King’s College London, Guy’s Hospital, London, United Kingdom

Botía J.A.:
 Reta Lila Weston Research Laboratories, Department of Molecular Neuroscience, University College London (UCL) Institute of Neurology, London, United Kingdom

 Departamento de Ingeniería de la Información y las Comunicaciones, Universidad de Murcia, Murcia, Spain

Vandrovcova J.:
 Reta Lila Weston Research Laboratories, Department of Molecular Neuroscience, University College London (UCL) Institute of Neurology, London, United Kingdom

Reynolds R.H.:
 Reta Lila Weston Research Laboratories, Department of Molecular Neuroscience, University College London (UCL) Institute of Neurology, London, United Kingdom

Zhang D.:
 Reta Lila Weston Research Laboratories, Department of Molecular Neuroscience, University College London (UCL) Institute of Neurology, London, United Kingdom

Trabzuni D.:
 Reta Lila Weston Research Laboratories, Department of Molecular Neuroscience, University College London (UCL) Institute of Neurology, London, United Kingdom

 Department of Genetics, King Faisal Specialist Hospital and Research Centre, Riyadh, Saudi Arabia

Collado-Torres L.:
 Lieber Institute for Brain Development, 855 North Wolfe Street, Baltimore, MD, United States

Thomason A.:
 Goldsmiths, University of London, New Cross, London, United Kingdom

Quijada Leyton P.:
 Goldsmiths, University of London, New Cross, London, United Kingdom

Gagliano Taliun S.A.:
 Center for Statistical Genetics, University of Michigan, Ann Arbor, MI, United States

Nalls M.A.:
 Laboratory of Neurogenetics, National Institute on Aging, US National Institutes of Health, Bethesda, MD, United States

 Data Tecnica International, Glen Echo, MD, United States

Noyce A.J.:
 Reta Lila Weston Research Laboratories, Department of Molecular Neuroscience, University College London (UCL) Institute of Neurology, London, United Kingdom

 Preventive Neurology Unit, Wolfson Institute of Preventive Medicine, QMUL, London, United Kingdom

Nicolas A.:
 Laboratory of Neurogenetics, National Institute on Aging, US National Institutes of Health, Bethesda, MD, United States

Cookson M.R.:
 Laboratory of Neurogenetics, National Institute on Aging, US National Institutes of Health, Bethesda, MD, United States

Bandres-Ciga S.:
 Laboratory of Neurogenetics, National Institute on Aging, US National Institutes of Health, Bethesda, MD, United States

Gibbs J.R.:
 Laboratory of Neurogenetics, National Institute on Aging, US National Institutes of Health, Bethesda, MD, United States

Hernandez D.G.:
 Laboratory of Neurogenetics, National Institute on Aging, US National Institutes of Health, Bethesda, MD, United States

Singleton A.B.:
 Laboratory of Neurogenetics, National Institute on Aging, US National Institutes of Health, Bethesda, MD, United States

Reed X.:
 Laboratory of Neurogenetics, National Institute on Aging, US National Institutes of Health, Bethesda, MD, United States

Leonard H.:
 Laboratory of Neurogenetics, National Institute on Aging, US National Institutes of Health, Bethesda, MD, United States

Blauwendraat C.:
 Laboratory of Neurogenetics, National Institute on Aging, US National Institutes of Health, Bethesda, MD, United States

 Preventive Neurology Unit, Wolfson Institute of Preventive Medicine, QMUL, London, United Kingdom

Faghri F.:
 Departamento de Ingeniería de la Información y las Comunicaciones, Universidad de Murcia, Murcia, Spain

 National Institute of Neurological Disorders and Stroke, Bethesda, MD, United States

 Department of Computer Science, University of Illinois at Urbana-Champaign, Urbana, IL, United States

Bras J.:
 Department of Twin Research and Genetic Epidemiology, King’s College London, London, United Kingdom

Guerreiro R.:
 Department of Molecular Neuroscience, UCL, London, United Kingdom

Tucci A.:
 Department of Molecular Neuroscience, UCL, London, United Kingdom

Kia D.A.:
 Department of Molecular Neuroscience, UCL, London, United Kingdom

Houlden H.:
 Department of Molecular Neuroscience, UCL, London, United Kingdom

Plun-Favreau H.:
 Department of Molecular Neuroscience, UCL, London, United Kingdom

Mok K.Y.:
 Department of Molecular Neuroscience, UCL, London, United Kingdom

Wood N.W.:
 Department of Molecular Neuroscience, UCL, London, United Kingdom

Lovering R.:
 Department of Molecular Neuroscience, UCL, London, United Kingdom

R’Bibo L.:
 Department of Molecular Neuroscience, UCL, London, United Kingdom

Rizig M.:
 Department of Molecular Neuroscience, UCL, London, United Kingdom

Chelban V.:
 Department of Molecular Neuroscience, UCL, London, United Kingdom

Tan M.:
 Singapore Institute for Clinical Sciences, Brenner Centre for Molecular Medicine, Singapore, Singapore

Morris H.R.:
 Department of Clinical Neuroscience, University College London, London, United Kingdom

Middlehurst B.:
 Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

Quinn J.:
 Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

Billingsley K.:
 Institute of Translational Medicine, University of Liverpool, Liverpool, United Kingdom

Holmans P.:
 Biostatistics & Bioinformatics Unit, Institute of Psychological Medicine and Clinical Neuroscience, MRC Centre for Neuropsychiatric Genetics & Genomics, Cardiff, United Kingdom

Kinghorn K.J.:
 Institute of Healthy Ageing, University College London, London, United Kingdom

Lewis P.:
 University of Reading, Reading, United Kingdom

Escott-Price V.:
 MRC Centre for Neuropsychiatric Genetics and Genomics, Cardiff University School of Medicine, Cardiff, United Kingdom

Williams N.:
 MRC Centre for Neuropsychiatric Genetics and Genomics, Cardiff University School of Medicine, Cardiff, United Kingdom

Foltynie T.:
 UCL Institute of Neurology, London, United Kingdom

Brice A.:
 Institut du Cerveau et de la Moelle épinière, ICM, Inserm U 1127, CNRS, UMR 7225, Sorbonne Universités, UPMC University Paris 06, UMR S 1127, AP-HP, Pitié-Salpêtrière Hospital, Paris, France

Danjou F.:
 Institut du Cerveau et de la Moelle épinière, ICM, Inserm U 1127, CNRS, UMR 7225, Sorbonne Universités, UPMC University Paris 06, UMR S 1127, AP-HP, Pitié-Salpêtrière Hospital, Paris, France

Lesage S.:
 Institut du Cerveau et de la Moelle épinière, ICM, Inserm U 1127, CNRS, UMR 7225, Sorbonne Universités, UPMC University Paris 06, UMR S 1127, AP-HP, Pitié-Salpêtrière Hospital, Paris, France

Corvol J.-C.:
 Institut du Cerveau et de la Moelle épinière, ICM, Inserm U 1127, CNRS, UMR 7225, Sorbonne Universités, UPMC University Paris 06, UMR S 1127, AP-HP, Pitié-Salpêtrière Hospital, Paris, France

Martinez M.:
 Institut du Cerveau et de la Moelle épinière, ICM, Inserm U 1127, CNRS, UMR 7225, Sorbonne Universités, UPMC University Paris 06, UMR S 1127, AP-HP, Pitié-Salpêtrière Hospital, Paris, France

 Paul Sabatier University, Toulouse, France

Giri A.:
 Department for Neurodegenerative Diseases, Hertie Institute for Clinical Brain Research, University of Tübingen, Tübingen, Germany

 DZNE, German Center for Neurodegenerative Diseases, Tübingen, Germany

Schulte C.:
 Department for Neurodegenerative Diseases, Hertie Institute for Clinical Brain Research, University of Tübingen, Tübingen, Germany

 DZNE, German Center for Neurodegenerative Diseases, Tübingen, Germany

Brockmann K.:
 Department for Neurodegenerative Diseases, Hertie Institute for Clinical Brain Research, University of Tübingen, Tübingen, Germany

 DZNE, German Center for Neurodegenerative Diseases, Tübingen, Germany

Simón-Sánchez J.:
 Department for Neurodegenerative Diseases, Hertie Institute for Clinical Brain Research, University of Tübingen, Tübingen, Germany

 DZNE, German Center for Neurodegenerative Diseases, Tübingen, Germany

Heutink P.:
 Department for Neurodegenerative Diseases, Hertie Institute for Clinical Brain Research, University of Tübingen, Tübingen, Germany

 DZNE, German Center for Neurodegenerative Diseases, Tübingen, Germany

Gasser T.:
 Department for Neurodegenerative Diseases, Hertie Institute for Clinical Brain Research, University of Tübingen, Tübingen, Germany

 DZNE, German Center for Neurodegenerative Diseases, Tübingen, Germany

Rizzu P.:
 DZNE, German Center for Neurodegenerative Diseases, Tübingen, Germany

Sharma M.:
 Centre for Genetic Epidemiology, Institute for Clinical Epidemiology and Applied Biometry, University of Tübingen, Tübingen, Germany

Shulman J.M.:
 Departments of Neurology, Neuroscience, and Molecular & Human Genetics, Baylor College of Medicine, Houston, TX, United States

 Jan and Dan Duncan Neurological Research Institute, Texas Children’s Hospital, Houston, TX, United States

Robak L.:
 Departments of Neurology, Neuroscience, and Molecular & Human Genetics, Baylor College of Medicine, Houston, TX, United States

Lubbe S.:
 Ken and Ruth Davee Department of Neurology, Northwestern University Feinberg School of Medicine, Chicago, IL, United States

 Northwestern University Feinberg School of Medicine, Chicago, IL, United States

Mencacci N.E.:
 Northwestern University Feinberg School of Medicine, Chicago, IL, United States

Finkbeiner S.:
 Departments of Neurology and Physiology, University of California, San Francisco, CA, United States

 Gladstone Institute of Neurological Disease

Lungu C.:
 National Institutes of Health Division of Clinical Research, NINDS, National Institutes of Health, Bethesda, MD, United States

Scholz S.W.:
 National Institute of Neurological Disorders and Stroke, Bethesda, MD, United States

 Neurodegenerative Diseases Research Unit, National Institute of Neurological Disorders and Stroke, Bethesda, MD, United States

Gan-Or Z.:
 Montreal Neurological Institute and Hospital, Department of Neurology & Neurosurgery, Department of Human Genetics, McGill University, Montréal, QC H3A 0G4, Canada

 Department of Human Genetics, McGill University, Montréal, QC H3A 0G4, Canada

Rouleau G.A.:
 Department of Human Genetics, McGill University, Montréal, QC H3A 0G4, Canada

Krohan L.:
 Department of Neurology, Leiden University Medical Center, Leiden, Netherlands

van Hilten J.J.:
 Instituto de Biomedicina de Sevilla (IBiS), Hospital Universitario Virgen del Rocío/CSIC/Universidad de Sevilla, Seville, Spain

Marinus J.:
 Instituto de Biomedicina de Sevilla (IBiS), Hospital Universitario Virgen del Rocío/CSIC/Universidad de Sevilla, Seville, Spain

Adarmes-Gómez A.D.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Bernal-Bernal I.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Bonilla-Toribio M.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Buiza-Rueda D.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Carrillo F.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Carrión-Claro M.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Mir P.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Gómez-Garre P.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Jesús S.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Labrador-Espinosa M.A.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Macias D.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Vargas-González L.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Méndez-del-Barrio C.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Periñán-Tocino T.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Tejera-Parrado C.:
 Fundació Docència i Recerca Mútua de Terrassa and Movement Disorders Unit, Department of Neurology, University Hospital Mutua de Terrassa, Terrassa, Barcelona, Spain

Diez-Fairen M.:
 Hospital Universitario Central de Asturias, Oviedo, Spain

Aguilar M.:
 Hospital Universitario Central de Asturias, Oviedo, Spain

Alvarez I.:
 Hospital Universitario Central de Asturias, Oviedo, Spain

Boungiorno M.T.:
 Hospital Universitario Central de Asturias, Oviedo, Spain

Carcel M.:
 Hospital Universitario Central de Asturias, Oviedo, Spain

Pastor P.:
 Hospital Universitario Central de Asturias, Oviedo, Spain

Tartari J.P.:
 Hospital Universitario Central de Asturias, Oviedo, Spain

Alvarez V.:
 Hospital Universitario Parque Tecnologico de la Salud, Granada, Spain

González M.M.:
 Hospital Universitario Parque Tecnologico de la Salud, Granada, Spain

Blazquez M.:
 Hospital Universitario Parque Tecnologico de la Salud, Granada, Spain

Garcia C.:
 Hospital Universitario Parque Tecnologico de la Salud, Granada, Spain

Suarez-Sanmartin E.:
 Hospital Universitario Parque Tecnologico de la Salud, Granada, Spain

Barrero F.J.:
 Instituto de Investigación Sanitaria Biodonostia, San Sebastián, Spain

Rezola E.M.:
 Hospital General de Segovia, Segovia, Spain

Yarza J.A.B.:
 Memory Unit, Department of Neurology, IIB Sant Pau, Hospital de la Santa Creu i Sant Pau, Universitat Autònoma de Barcelona, Barcelona, Spain

Pagola A.G.:
 Memory Unit, Department of Neurology, IIB Sant Pau, Hospital de la Santa Creu i Sant Pau, Universitat Autònoma de Barcelona, Barcelona, Spain

Arregui A.L.M.:
 Memory Unit, Department of Neurology, IIB Sant Pau, Hospital de la Santa Creu i Sant Pau, Universitat Autònoma de Barcelona, Barcelona, Spain

Ruiz-Martínez J.:
 Memory Unit, Department of Neurology, IIB Sant Pau, Hospital de la Santa Creu i Sant Pau, Universitat Autònoma de Barcelona, Barcelona, Spain

Cerdan D.:
 Centro de Investigación Biomédica en Red en Enfermedades Neurodegenerativas (CIBERNED), Madrid, Spain

Duarte J.:
 Hospital Universitario Marqués de Valdecilla-IDIVAL, Santander, Spain

Clarimón J.:
 University of Cantabria, Santander, Spain

 Movement Disorders Unit, Department of Neurology, IIB Sant Pau, Hospital de la Santa Creu i Sant Pau, Universitat Autònoma de Barcelona, Barcelona, Spain

Dols-Icardo O.:
 University of Cantabria, Santander, Spain

 Movement Disorders Unit, Department of Neurology, IIB Sant Pau, Hospital de la Santa Creu i Sant Pau, Universitat Autònoma de Barcelona, Barcelona, Spain

Infante J.:
 Movement Disorders Unit, Department of Neurology, IIB Sant Pau, Hospital de la Santa Creu i Sant Pau, Universitat Autònoma de Barcelona, Barcelona, Spain

 Centro de Investigacion Biomedica, Universidad de Granada, Granada, Spain

 Hospital Universitario Virgen de las Nieves, Instituto de Investigación Biosanitaria de Granada, Granada, Spain

Marín J.:
 Movement Disorders Unit, Department of Neurology, IIB Sant Pau, Hospital de la Santa Creu i Sant Pau, Universitat Autònoma de Barcelona, Barcelona, Spain

 Hospital Clínic de Barcelona, Barcelona, Spain

Kulisevsky J.:
 Movement Disorders Unit, Department of Neurology, IIB Sant Pau, Hospital de la Santa Creu i Sant Pau, Universitat Autònoma de Barcelona, Barcelona, Spain

 Hospital Clínic de Barcelona, Barcelona, Spain

Pagonabarraga J.:
 Movement Disorders Unit, Department of Neurology, IIB Sant Pau, Hospital de la Santa Creu i Sant Pau, Universitat Autònoma de Barcelona, Barcelona, Spain

 Hospital Clínic de Barcelona, Barcelona, Spain

Gonzalez-Aramburu I.:
 Centro de Investigacion Biomedica, Universidad de Granada, Granada, Spain

Rodriguez A.S.:
 Centro de Investigacion Biomedica, Universidad de Granada, Granada, Spain

Sierra M.:
 Centro de Investigacion Biomedica, Universidad de Granada, Granada, Spain

Duran R.:
 Instituto de Investigación Sanitaria Fundación Jiménez Díaz, Madrid, Spain

Ruz C.:
 Instituto de Investigación Sanitaria Fundación Jiménez Díaz, Madrid, Spain

Vives F.:
 Instituto de Investigación Sanitaria Fundación Jiménez Díaz, Madrid, Spain

Escamilla-Sevilla F.:
 Hospital Universitario Virgen de la Victoria, Málaga, Spain

Mínguez A.:
 Hospital Universitario Virgen de la Victoria, Málaga, Spain

Cámara A.:
 Institut de Recerca Sant Joan de Déu, Barcelona, Spain

Compta Y.:
 Institut de Recerca Sant Joan de Déu, Barcelona, Spain

Ezquerra M.:
 Institut de Recerca Sant Joan de Déu, Barcelona, Spain

Marti M.J.:
 Institut de Recerca Sant Joan de Déu, Barcelona, Spain

Fernández M.:
 Institut de Recerca Sant Joan de Déu, Barcelona, Spain

Muñoz E.:
 Institut de Recerca Sant Joan de Déu, Barcelona, Spain

Fernández-Santiago R.:
 Institut de Recerca Sant Joan de Déu, Barcelona, Spain

Tolosa E.:
 Institut de Recerca Sant Joan de Déu, Barcelona, Spain

Valldeoriola F.:
 Institut de Recerca Sant Joan de Déu, Barcelona, Spain

García-Ruiz P.:
 Hospital Universitario Ramón y Cajal, Madrid, Spain

Heredia M.J.G.:
 Department of Neurology, Instituto de Investigación Sanitaria La Fe, Hospital Universitario y Politécnico La Fe, Valencia, Spain

Errazquin F.P.:
 Department of Neurology, Instituto de Investigación Sanitaria La Fe, Hospital Universitario y Politécnico La Fe, Valencia, Spain

Hoenicka J.:
 Hospital General de Segovia, Segovia, Spain

Jimenez-Escrig A.:
 Hospital General de Segovia, Segovia, Spain

Martínez-Castrillo J.C.:
 Hospital General de Segovia, Segovia, Spain

Lopez-Sendon J.L.:
 Hospital General de Segovia, Segovia, Spain

Torres I.M.:
 Department of Neurology, Hospital Universitario Fundación Alcorcón, Madrid, Spain

Tabernero C.:
 Department of Neurology, Hospital Universitario Fundación Alcorcón, Madrid, Spain

Vela L.:
 Department of Neurology, Hospital Universitario Fundación Alcorcón, Madrid, Spain

Zimprich A.:
 Department of Neurology, Medical University of Vienna, Vienna, Austria

Pihlstrom L.:
 Department of Neurology, Oslo University Hospital, Oslo, Norway

Koks S.:
 Department of Pathophysiology, University of Tartu, Tartu, Estonia

 Department of Reproductive Biology, Estonian University of Life Sciences, Tartu, Estonia

 Perron Institute for Neurological and Translational Science, Perth, WA, Australia

Taba P.:
 Department of Neurology and Neurosurgery, University of Tartu, Tartu, Estonia

Majamaa K.:
 Institute of Clinical Medicine, Department of Neurology, University of Oulu, Oulu, Finland

 Department of Neurology and Medical Research Center, Oulu University Hospital, Oulu, Finland

Siitonen A.:
 Institute of Clinical Medicine, Department of Neurology, University of Oulu, Oulu, Finland

 Department of Neurology and Medical Research Center, Oulu University Hospital, Oulu, Finland

Okubadejo N.U.:
 University of Lago, Lagos State, Nigeria

Ojo O.O.:
 University of Lago, Lagos State, Nigeria

Forabosco P.:
 Istituto di Ricerca Genetica e Biomedica, Cittadella Universitaria di Cagliari, Monserrato, Sardinia 09042, Italy

Walker R.:
 Department of Neuropathology, MRC Sudden Death Brain Bank Project, University of Edinburgh, Edinburgh, United Kingdom

Small K.S.:
 Department of Twin Research and Genetic Epidemiology, King’s College London, London, United Kingdom

Smith C.:
 Department of Neuropathology, MRC Sudden Death Brain Bank Project, University of Edinburgh, Edinburgh, United Kingdom

Ramasamy A.:
 Reta Lila Weston Research Laboratories, Department of Molecular Neuroscience, University College London (UCL) Institute of Neurology, London, United Kingdom

 Department of Medical & Molecular Genetics, School of Medical Sciences, King’s College London, Guy’s Hospital, London, United Kingdom

 Singapore Institute for Clinical Sciences, Brenner Centre for Molecular Medicine, Singapore, Singapore

Hardy J.:
 Reta Lila Weston Research Laboratories, Department of Molecular Neuroscience, University College London (UCL) Institute of Neurology, London, United Kingdom

Weale M.E.:
 Department of Medical & Molecular Genetics, School of Medical Sciences, King’s College London, Guy’s Hospital, London, United Kingdom

 Genomics PLC, Oxford, United Kingdom

Ryten M.:
 Reta Lila Weston Research Laboratories, Department of Molecular Neuroscience, University College London (UCL) Institute of Neurology, London, United Kingdom
ISSN: 20411723
Editorial
NATURE PUBLISHING GROUP, MACMILLAN BUILDING, 4 CRINAN ST, LONDON N1 9XW, ENGLAND, Reino Unido
Tipo de documento: Article
Volumen: 11 Número: 1
Páginas:
WOS Id: 000564282900001
ID de PubMed: 32098967
imagen Green Published, gold, Gold, Green

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